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    • Animals - Kingdom: Animalia >
      • Thorny-headed Worms - Phylum: Acanthocephala
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            • True Yams - Family: Dioscoreaceae
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              • Sunflowers, Asters, Daisies and Allies (Part 4.) - Family: Asteraceae
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Wolf  Spiders  and  Allies  -  Superfamily:   Lycosoidea

      The Lycosoidea as a superfamily typically consisting of families of spiders that share a particular feature of their eyes, namely the shape of the tapetum (= a reflective layer at the back of some or all of the eyes, believed to enhance light sensitivity).  In Lycosoids this tapetum is supposedly 'grate-shaped'.  As a defining feature this has not always been satisfactory however.  and a number of revisions have taken place that have seen families added, and a number of species and genera re-arranged.  The current taxonomic configuration is still debated and, new discoveries are still being made, further complicating the matter.
      The Lycosoid spiders are mostly vagrants, meaning they do not build webs to catch their prey but, rather, they either actively hunt for it or they ambush it using camouflage and stealth.  As hunting spiders many do not shy away from tackling prey larger than themselves, some are quite adept at catching small fish and others are aggressive enough to attack and kill other spiders for a meal.  They can be found on every continent except Antarctica although some species have been sighted well above the Arctic circle.   Many species are rather drab looking but there are plenty of exceptions in most Lycosoid families.

      This superfamily is composed of 10 families, altogether accounting for 311 genera and 4,420 species as of November 2023.   The largest family is the Lycosidae with about 55 % of the total species count.  Six of the ten families are represented in the Northern Rivers.
Lycosoid spiders may have appeared during the Cretaceous Period as suggested by a recent find in Botswana of what is regarded to be a fossil Lycosoid spider dated some 113-101 My old.  The fossil spider is said to be 'almost-complete' and is the first such specimen to be described from Africa.

Wandering Spiders Family: Ctenidae
Wolf Spiders - Family: Lycosidae
Lynx Spiders - Family: Oxyopidae
Nursery Web Spiders Family: Pisauridae
Sheetweb spiders - Family: Stiphidiidae
False Wolf Spiders Family: Zoropsidae


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References  and  links:
​

>      Benson, K.,  and  Suter, R.B.,   (2013).    'Reflections on the tapetum lucidum and eyeshine in lycosoid spiders.'    The Journal of Arachnology. 41 (1): 43–52. doi:10.1636/B12-35.1.
>      Carico, J.E.,  (2005).   'Descriptions of two new spider genera of Trechaleidae (Araneae, Lycosoidea) from South America.'   Journal of Arachnology 33(3): 797-812. 
​
doi:10.1636/H03-71.1

​>      Gray, M.R.  and  Thompson, J.A.,  (2001).    'New lycosoid spiders from cave and surface habitats in southern Australia and Cape Range peninsula (Araneae: Lycosoidea).'   Records of the Western Australian Museum, Supplement 64: 159-170.   doi:10.18195/issn.0313-122x.64.2001.159-170 
>      Griswold, C E.,  (1993).   'Investigations into the phylogeny of Lycosoid spiders and their kin (Arachnida: Araneae: Lycosoidea).'    Smithsonian Contributions to Zoology. 539 (539):
​ 1–39.   
doi:10.5479/si.00810282.539

>      Polotow, D.,  Carmichael, A.  and  Griswold, C.E.,   (2015).    'Total evidence analysis of the phylogenetic relationships of Lycosoidea spiders (Araneae, Entelegynae).'   Invertebrate Systematics 29(2): 124-163. doi:10.1071/IS14041
>      Rayner, R.J. and Dippenaar-Schoeman, A.S.,   (1995).    'A fossil spider (superfamily Lycosoidea) from the Cretaceous of Botswana.'    South African Journal of Science 91, 98–100.
>      Selden, P.,  Nam, K.-S.,   Kim, S.-H.  and  Kim, Hyun,  (2012).    'A Fossil Spider from the Cretaceous of Korea.'   Journal of Paleontology. 86. 1-6.   doi:10.2307/41409126. 

      https://en.wikipedia.org/wiki/Lycosoidea
      https://inaturalist.ala.org.au/taxa/367161-Lycosoidea
      https://www.mindat.org/taxon-P57517.html


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  • Home
    • About the Northern Rivers Project website, and, how to use it.
  • Life
    • Easy navigator to the LIFE pages. >
      • Arachnids, Spiders, Harvestmen, Scorpions, Ticks and Mites of the Northern Rivers, Index
      • Birds, Index
      • Butterflies of the Northern Rivers region of New South Wales, Index
      • Mammals of the Northern Rivers region of New South Wales, Index
      • Reptiles of the Northern Rivers region of New South Wales, Index
    • Animals - Kingdom: Animalia >
      • Thorny-headed Worms - Phylum: Acanthocephala
      • Segmented Worms - Phylum: Annelida
      • Arthropods - Phylum: Arthropoda
      • Brachiopods - Phylum Brachiopoda
      • Bryozoans - Phylum: Bryozoa
      • Arrow Worms - Phylum: Chaetognatha
      • Chordates - Phylum: Chordata
      • Cnidarians - Phylum: Cnidaria
      • Cycliophorans - Phylum: Cycliophora
      • Dicyemids - Phylum: Dicyemida
      • Echinoderms - Phylum: Echinodermata
      • Entoprocts - Phylum: Entoprocta
      • Gastrotrichs - Phylum: Gastrotricha
      • Gnathiferans - Phylum: Gnathifera
      • Jaw Worms - Phylum: Gnathostomulida
      • Hemichordates - Phylum: Hemichordata
      • Mud Dragons - Phylum: Kinorhyncha
      • Loriciferans - Phylum: Loricifera
      • Molluscs- Phylum: Mollusca
      • Nematodes - Phylum: Nematoda
      • Horsehair Worms - Phylum: Nematomorpha
      • Ribbon Worms - Phylum: Nemertea
      • Velvet Worms - Phylum: Onychophora >
        • Modern Velvet Worms - Order: Euonychophora >
          • Southern Velvet Worms - Family: Peripatopsidae
      • Orthonectids - Phylum: Orthonectida
      • Horseshoe Worms - Phylum: Phoronida
      • Placozoans - Phylum: Placozoa
      • Flatworms - Phylum: Platyhelminthes >
        • Rhabditophora (Subphylum) >
          • Planarians - Order: Tricladida >
            • Continenticola (Suborder) >
              • Geoplanoidea (Superfamily) >
                • Land Planarians - Family: Geoplanidae
      • Sponges - Phylum: Porifera
      • Priapulid Worms - Phylum: Priapulida
      • Rotifers - Phylum: Rotifera
      • Tardigrades - Phylum: Tardigrada
      • Xenoturbellids - Phylum: Xenacoelomorpha
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        • True Mosses - Class: Bryopsida >
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      • Charophyte Algae - Phylum: Charophyta
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      • Liverworts - Phylum: Marchantiophyta
      • Red Algae - Phylum: Rhodophyta
      • Vascular Plants - Phylum: Tracheophyta >
        • Cycads and Zamias - Class: Cycadopsida >
          • Zamias - Family: Zamiaceae
        • Lycophytes - Class: Lycopodiopsida >
          • Clubmosses - Order: Lycopodiales >
            • Clubmosses - Family: Lycopodiaceae
        • Conifers - Class: Pinopsida >
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        • Ferns - Class: Polypodiopsida >
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            • Tree Ferns - Order: Cyatheales >
              • Scaly Tree Ferns - Family: Cyatheaceae
              • Hairy Tree Ferns - Family: Dicksoniaceae
            • Forked Ferns - Order: Gleicheniales >
              • Forked Ferns - Family: Gleicheniaceae
            • Filmy Ferns - Order: Hymenophyllales >
              • Filmy Ferns - Family: Hymenophyllaceae
            • Polypod Ferns - Order: Polypodiales >
              • Eupolypods ll - Suborder: Aspleniineae >
                • Spleenworts - Family: Aspleniaceae
                • Chain Ferns - Family: Blechnaceae
              • Brackens - Suborder: Dennstaedtiineae >
                • Brackens - Family: Dennstaedtiaceae
              • Eupolypods l - Suborder: Polypodiineae >
                • Sword Ferns - Family: Nephrolepidaceae
                • Polypod Ferns - Family: Polypodiaceae
                • Tectarias - Family: Tectariaceae
              • Brakes - Suborder: Pteridineae >
                • Brakes - Family: Pteridaceae
            • Aquatic Ferns - Order: Salviniales >
              • Water Ferns - Family: Salviniaceae
        • Monocotyledons - Class: Liliopsida >
          • Water Plantains and Allies - Order: Alismatales >
            • Water Plantains - Family: Alismataceae
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            • Palms - Family: Arecaceae
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            • Amaryllis, Onions and Allies - Family: Amaryllidaceae
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              • Orchids (Part 6.) - Family: Orchidaceae
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            • Spiderworts - Family: Commelinaceae
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          • Yams, Colicroots and Allies - Order: Dioscoreales >
            • True Yams - Family: Dioscoreaceae
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            • Bellworts, Naked Ladies and Allies - Family: Colchicaceae
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            • Screwpines - Family: Pandanaceae
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            • Arrowroots - Family: Marantaceae
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        • Dicotyledons - Class: Magnoliopsida >
          • Carrots, Ivies and Allies - Order: Apiales >
            • Umbellifers - Family: Apiaceae
            • Ivies and Allies - Family: Araliaceae
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          • Hollies and Allies - Order: Aquifoliales >
            • Cardiopteris - Family: Cardiopteridaceae
          • Sunflowers, Bellflowers, Fanflowers and Allies - Order: Asterales >
            • Korokios and Allies - Family: Argyophyllaceae
            • Sunflowers, Asters, Daisies and Allies - Family: Asteraceae >
              • Sunflowers, Asters, Daisies and Allies (Part 1.) - Family: Asteraceae
              • Sunflowers, Asters, Daisies and Allies (Part 2.) - Family: Asteraceae
              • Sunflowers, Asters, Daisies and Allies (Part 3.) - Family: Asteraceae
              • Sunflowers, Asters, Daisies and Allies (Part 4.) - Family: Asteraceae
            • Bellflowers, Lobelias and Allies - Family: Campanulaceae
            • Fanflowers, Goodenias and Allies - Family: Goodeniaceae
            • Bogbeans and Allies - Family: Menyanthaceae
          • Borage and Allies - Order: Boraginales >
            • Borage - Family: Boraginaceae
          • Mustards, Capers and Allies - Ordser: Brassicales >
            • Mustards and Allies - Family: Brassicaceae
            • Capers and Allies - Family: Capparaceae
            • Papayas - Family: Caricaceae
          • Canellas, Winter's Bark and Allies - Order: Canellales
          • Pinks, Cactuses and Allies - Order: Caryophyllales >
            • Stone Plants - Family: Aizoaceae
            • Amaranths and Allies - Family: Amaranthaceae
            • Basellas - Family: Basellaceae
            • Cactuses - Family: Cactaceae
            • Pinks, Chickweeds and Allies - Family: Caryophyllaceae
            • Sundews - Family: Droseraceae
            • Umbrellaworts - Family: Nyctaginaceae
            • Pokeweeds - Family: Phytolaccaceae
            • Knotweeds - Family: Polygonaceae
            • Purslanes - Family: Portulaceae
          • Staff-vines and Allies - Order: Celastrales >
            • Staff-vines - Family: Celastraceae
          • Gourds, Begonias and Allies - Order: Cucurbitales
          • Guinea Flowers and Allies - Order: Dilleniales >
            • Guinea Flowers - Family: Dilleniaceae
          • Heaths, Balsams, Primroses and Allies - Order: Ericales >
            • Balsams and Allies - Family: Balsaminaceae
            • Persimmons and Allies - Family: Ebenaceae
            • Heaths - Family: Ericaceae
            • Primroses - Family: Primulaceae
            • Sapodillas - Family: Sapotaceae
          • Escallonias and Allies - Order: Escalloniales >
            • Escallonias - Family: Escalloniaceae
          • Legumes, Milkworts and Allies - Order: Fabales >
            • Legumes - Family: Fabaceae >
              • Legumes (Part 1) - Family: Fabaceae
              • Legumes (Part 2) - Family: Fabaceae
              • Legumes (Part 3) - Family: Fabaceae
          • Beeches, Oaks, Walnuts and Allies - Order: Fagales >
            • Birches and Allies - Family: Betulaceae
            • Sheoaks - Family: Casuarinaceae
            • Oaks - Family: Fagaceae
            • Southern Beeches - Family: Nothofagaceae
            • Walnuts - Family: Juglandaceae
          • Gentians, Dogbanes, Madders and Allies - Order: Gentianales >
            • Dogbanes - Family: Apocynaceae
            • Madders - Family: Rubiaceae
          • Geraniums, Cranesbills and Allies - Order: Geraniales >
            • Geraniums and Cranesbills - Family: Geraniaceae
          • Mints, Plantains, Olives and Allies - Order: Lamiales >
            • Acanthuses and Allies - Family: Acanthaceae
            • Bignonias - Family: Bignoniaceae
            • Gesnerias - Family: Gesneriaceae
            • Mints - Family: Lamiaceae
            • Bladderworts - Family: Lentibulariaceae
            • Olives and Allies - Family: Oleaceae
            • Plantains - Family: Plantaginaceae
            • Figworts - Family: Scrophulariaceae
            • Verbenas - Family: Verbenaceae
          • Laurels, Spicebushes and Allies - Order: Laurales >
            • Laurels - Family: Lauraceae
          • Magnolias, Nutmegs and Allies - Order: Magnoliales >
            • Bolwarras - Family: Eupomatiaceae
          • Nances, Willows and Allies - Order: Malpighiales >
            • Spurges - Family: Euphorbiaceae
            • Ochnas - Family: Ochnaceae
            • Passionflowers - Family: Passifloraceae
            • Leaf-flowers - Family: Phyllanthaceae
            • Picrodendrons - Family: Picrodendraceae
            • Ironplums - Family: Putranjivaceae
            • Mangroves - Family: Rhizoporaceae
            • Violets - Family: Violaceae
          • Mallows, Rock-roses and Allies - Order: Malvales >
            • Stringbarks - Family: Thymelaeaceae
            • Mallows, Hibiscuses and Allies - Family: Malvaceae
          • Myrtles, Evening Primroses and Allies - Order: Myrtales >
            • Lythrums and Allies - Family: Lythraceae
            • Melastomes - Family: Melastomataceae
            • Myrtles, Eucalypts, Melaleucas and Allies - Family: Myrtaceae >
              • Waxflowers and Allies - Tribe: Chamelaucieae
              • Gum Trees - Tribe: Eucalypteae
              • Tea Trees and Myrtles - Tribe: Leptospermeae
              • Melaleucas - Tribe: Melaleuceae
              • Waterberries - Tribe: Syzygieae
              • All other Tribes - Subfamily: Myrtoidea
            • Evening Primroses and Allies - Family: Onagraceae
          • Waterlilies, Fanworts and Allies - Order: Nymphaeales >
            • Waterlilies - Family: Nymphaeaceae
          • Woodsorrels, Quandongs and Allies - Order: Oxalidales >
            • Butterspoons - Family: Cunoniaceae
            • Quandongs - Family: Elaecarpaceae
            • Woodsorrels - Family: Oxalidaceae
          • Pepper Plants, Birthworts and Allies - Order: Piperales >
            • Pepper Plants - Family: Piperaceae
          • Proteas, Lotuses, Planes and Allies - Order: Proteales >
            • Proteas and Allies - Family: Proteaceae >
              • Grevilleas, Banksias and Allies - Subfamily: Grevilleoidea >
                • Banksias - Tribe: Banksieae
                • Firebushes and Allies - Tribe: Embothrieae
                • Macadamias and Allies - Tribe: Macadamieae
                • Honeysuckles and Allies - Tribe: Roupaleae
              • Primitive Proteas - Subfamily: Persoonioideae >
                • Geebungs and Allies - Tribe: Persoonieae
              • Singlet Proteas - Subfamily: Proteoideae >
                • Featherbushes and Conesticks - Tribe: Petrophileae
          • Buttercups, Poppies and Allies - Order: Ranunculales >
            • Buttercups - Family: Ranunculaceae
            • Moonseeds - Family: Menispermaceae
            • Poppies and Allies - Family: Papaveraceae
          • Roses, Figs, Elms and Allies - Order: Rosales >
            • Hemps and Allies - Family: Cannabaceae
            • Figs, Mulberries and Allies - Family: Moraceae
            • Buckthorns and Allies - Family: Rhamnaceae
            • Roses, Brambles and Allies - Family: Rosaceae
            • Elms and Allies - Family: Ulmaceae
            • Nettles and Allies - Family: Urticaceae
          • Sandalwoods, Mistletoes and Allies - Order: Santalales >
            • Showy Mistletoes - Family: Loranthaceae
            • Sandalwoods - Family: Santalaceae
          • Soapberries, Mahoganies and Allies - Order: Sapindales >
            • Cashews, Sumacs and Allies - Family: Anacardiaceae
            • Mahoganies and Allies - Family: Meliaceae
            • Rues, Citruses and Allies - Family: Rutaceae
            • Soapberries and Allies - Family: Sapindaceae
            • Quassias, Corkwoods and Allies - Family: Simaroubaceae
          • Saxifrages, Stonecrops and Allies - Order: Saxifragales >
            • Stonecrops and Allies - Family: Crassulaceae
          • Nightshades, Bindweeds and Allies - Order: Solanales >
            • Bindweeds, Ponysfoots and Allies - Family: Convolvulaceae
            • Nightshades, Tobacco and Allies - Family: Solanaceae
          • Grapes and Allies - Order: Vitales >
            • Grapes and Allies - Family: Vitaceae
    • Protozoans - Kingdom: Protozoa >
      • Slime Moulds - Phylum: Mycetozoa >
        • True Slime Moulds - Class: Myxomycetes
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