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Open-holed  Trapdoor  Spiders  -  Family:  Pycnothelidae

​​      The Pycnothelidae are a family of 139 species in 15 genera. It should be borne in mind though that over the past 3 to 4 years the family has been the subject of a number of phylogenetic studies, having been re-established as a family in its' own right (2020) after R. Raven, in 1985, downgraded it to the status of subfamily in the Nemesiidae.  It is currently accepted to contain seven subfamilies as mentioned on the 'World Spider Catalog'.    The majority of species are endemic to South America, a couple of small genera to Africa and one genus, Stanwellia, contains species species endemic to New Zealand or Australia only.  The Genus: Stanwellia was first described by Rainbow & Pulleine in 1918.
      For the eight Stanwellia species found in Australia most records have been from the south-east of the country with Victoria the state where they are most common.  There have only been a handful of records of Stanwellia spp. in the Northern Rivers although the sightings have not been identified to species.   Stanwellia species do bear quite a resemblance to mygalomorphs of the Genus: Aname (fam: Anamidae) though and even if not likely, the possibility of misidentification can not be excluded.  For now, the Pycnothelidae are included here until such time as it is proven to be inappropriate.

      Open-holed Trapdoor Spiders from the Genus: Stanwellia are generally large spiders, of the usual colour palette that dominates all mygalomorphs.  They are robustly buiilt with strong, shortish legs.  Two of their spinnerets protrude from underneath the abdomen like two little tails.  As the name suggests they do build burrows, with a silk lining but normally without a lid. These burrows may be up to 40 cm deep.


Subfamily:  Diplothelopsinae

      Stanwellia sp.

​      


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Taxonomy:
​​
Phylum:   Arthropoda   (Arthropods)
          > Subphylum:   Chelicerata   (Chelicerates)
                    > Class:   Arachnida   (Arachnids) 
                                        > Order:   Aranea   (Spiders)
                                                  > Suborder:   Mygalomorphae   (Mygalomorphs)

                                                                      > Superfamily:   Avicularioidea
                                                                                > Family:   Pycnothelidae   (Open-holded Trapdoor Spiders)
                                                                                          > Subfamily:   Diplothelopsinae
                                                                                                              > Genus:   Stanwellia 
 ^  (Open-holded Trapdoor Spiders)
                                                                                                                        > Species:  ?

*  =  likely
^  =  possibly

References  and  links:
​
>      Montes de Oca, L.,  Indicatti, R.P.,  Opatova, V.,  Almeida, M.,  Pérez-Miles, F.   and  Bond, J.E.,  (2022).   'Phylogenomic analysis, reclassification, and evolution of South American nemesioid burrowing mygalomorph spiders.'   Molecular Phylogenetics and Evolution 168(107377): 1-19.   doi:10.1016/j.ympev.2021.107377
>      Rainbow, W.J.   and  Pulleine, R.H.,  (1918).    'Australian trap-door spiders.'   Records of the Australian Museum 12: 81-169, pl. 12-24.

      https://wsc.nmbe.ch/family/138/Pycnothelidae
      https://www.arachne.org.au/01_cms/details.asp?ID=2686
​

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            • Spiderworts - Family: Commelinaceae
            • Stream Lilies and Frog-mouths - Family: Phylidraceae
          • Yams, Colicroots and Allies - Order: Dioscoreales >
            • True Yams - Family: Dioscoreaceae
          • Lilies, Supplejacks and Allies - Order: Liliales >
            • Bellworts, Naked Ladies and Allies - Family: Colchicaceae
            • Greenbriers - Family: Smilacaceae
          • Screwpines and Allies - Order: Pandanales >
            • Screwpines - Family: Pandanaceae
          • Grasses and Sedges - Order: Poales >
            • Bromeliads - Family: Bromeliaceae
            • Sedges - Family: Cyperaceae
            • Grasses - Family: Poaceae
            • Restiads - Family: Restionaceae
            • Bulrushes, Cattails and Allies - Family: Typhaceae
          • Canna Lilies and Gingers - Order: Zingiberales >
            • Arrowroots - Family: Marantaceae
            • Gingers - Family: Zingiberaceae
        • Dicotyledons - Class: Magnoliopsida >
          • Carrots, Ivies and Allies - Order: Apiales >
            • Umbellifers - Family: Apiaceae
            • Ivies and Allies - Family: Araliaceae
            • Cheesewoods and Allies - Family: Pittosporaceae
          • Hollies and Allies - Order: Aquifoliales >
            • Cardiopteris - Family: Cardiopteridaceae
          • Sunflowers, Bellflowers, Fanflowers and Allies - Order: Asterales >
            • Korokios and Allies - Family: Argyophyllaceae
            • Sunflowers, Asters, Daisies and Allies - Family: Asteraceae >
              • Sunflowers, Asters, Daisies and Allies (Part 1.) - Family: Asteraceae
              • Sunflowers, Asters, Daisies and Allies (Part 2.) - Family: Asteraceae
              • Sunflowers, Asters, Daisies and Allies (Part 3.) - Family: Asteraceae
              • Sunflowers, Asters, Daisies and Allies (Part 4.) - Family: Asteraceae
            • Bellflowers, Lobelias and Allies - Family: Campanulaceae
            • Fanflowers, Goodenias and Allies - Family: Goodeniaceae
            • Bogbeans and Allies - Family: Menyanthaceae
          • Borage and Allies - Order: Boraginales >
            • Borage - Family: Boraginaceae
          • Mustards, Capers and Allies - Ordser: Brassicales >
            • Mustards and Allies - Family: Brassicaceae
            • Capers and Allies - Family: Capparaceae
            • Papayas - Family: Caricaceae
          • Canellas, Winter's Bark and Allies - Order: Canellales
          • Pinks, Cactuses and Allies - Order: Caryophyllales >
            • Stone Plants - Family: Aizoaceae
            • Amaranths and Allies - Family: Amaranthaceae
            • Basellas - Family: Basellaceae
            • Cactuses - Family: Cactaceae
            • Pinks, Chickweeds and Allies - Family: Caryophyllaceae
            • Sundews - Family: Droseraceae
            • Umbrellaworts - Family: Nyctaginaceae
            • Pokeweeds - Family: Phytolaccaceae
            • Knotweeds - Family: Polygonaceae
            • Purslanes - Family: Portulaceae
          • Staff-vines and Allies - Order: Celastrales >
            • Staff-vines - Family: Celastraceae
          • Gourds, Begonias and Allies - Order: Cucurbitales
          • Guinea Flowers and Allies - Order: Dilleniales >
            • Guinea Flowers - Family: Dilleniaceae
          • Heaths, Balsams, Primroses and Allies - Order: Ericales >
            • Balsams and Allies - Family: Balsaminaceae
            • Persimmons and Allies - Family: Ebenaceae
            • Heaths - Family: Ericaceae
            • Primroses - Family: Primulaceae
            • Sapodillas - Family: Sapotaceae
          • Escallonias and Allies - Order: Escalloniales >
            • Escallonias - Family: Escalloniaceae
          • Legumes, Milkworts and Allies - Order: Fabales >
            • Legumes - Family: Fabaceae >
              • Legumes (Part 1) - Family: Fabaceae
              • Legumes (Part 2) - Family: Fabaceae
              • Legumes (Part 3) - Family: Fabaceae
          • Beeches, Oaks, Walnuts and Allies - Order: Fagales >
            • Birches and Allies - Family: Betulaceae
            • Sheoaks - Family: Casuarinaceae
            • Oaks - Family: Fagaceae
            • Southern Beeches - Family: Nothofagaceae
            • Walnuts - Family: Juglandaceae
          • Gentians, Dogbanes, Madders and Allies - Order: Gentianales >
            • Dogbanes - Family: Apocynaceae
            • Madders - Family: Rubiaceae
          • Geraniums, Cranesbills and Allies - Order: Geraniales >
            • Geraniums and Cranesbills - Family: Geraniaceae
          • Mints, Plantains, Olives and Allies - Order: Lamiales >
            • Acanthuses and Allies - Family: Acanthaceae
            • Bignonias - Family: Bignoniaceae
            • Gesnerias - Family: Gesneriaceae
            • Mints - Family: Lamiaceae
            • Bladderworts - Family: Lentibulariaceae
            • Olives and Allies - Family: Oleaceae
            • Plantains - Family: Plantaginaceae
            • Figworts - Family: Scrophulariaceae
            • Verbenas - Family: Verbenaceae
          • Laurels, Spicebushes and Allies - Order: Laurales >
            • Laurels - Family: Lauraceae
          • Magnolias, Nutmegs and Allies - Order: Magnoliales >
            • Bolwarras - Family: Eupomatiaceae
          • Nances, Willows and Allies - Order: Malpighiales >
            • Spurges - Family: Euphorbiaceae
            • Ochnas - Family: Ochnaceae
            • Passionflowers - Family: Passifloraceae
            • Leaf-flowers - Family: Phyllanthaceae
            • Picrodendrons - Family: Picrodendraceae
            • Ironplums - Family: Putranjivaceae
            • Mangroves - Family: Rhizoporaceae
            • Violets - Family: Violaceae
          • Mallows, Rock-roses and Allies - Order: Malvales >
            • Stringbarks - Family: Thymelaeaceae
            • Mallows, Hibiscuses and Allies - Family: Malvaceae
          • Myrtles, Evening Primroses and Allies - Order: Myrtales >
            • Lythrums and Allies - Family: Lythraceae
            • Melastomes - Family: Melastomataceae
            • Myrtles, Eucalypts, Melaleucas and Allies - Family: Myrtaceae >
              • Waxflowers and Allies - Tribe: Chamelaucieae
              • Gum Trees - Tribe: Eucalypteae
              • Tea Trees and Myrtles - Tribe: Leptospermeae
              • Melaleucas - Tribe: Melaleuceae
              • Waterberries - Tribe: Syzygieae
              • All other Tribes - Subfamily: Myrtoidea
            • Evening Primroses and Allies - Family: Onagraceae
          • Waterlilies, Fanworts and Allies - Order: Nymphaeales >
            • Waterlilies - Family: Nymphaeaceae
          • Woodsorrels, Quandongs and Allies - Order: Oxalidales >
            • Butterspoons - Family: Cunoniaceae
            • Quandongs - Family: Elaecarpaceae
            • Woodsorrels - Family: Oxalidaceae
          • Pepper Plants, Birthworts and Allies - Order: Piperales >
            • Pepper Plants - Family: Piperaceae
          • Proteas, Lotuses, Planes and Allies - Order: Proteales >
            • Proteas and Allies - Family: Proteaceae >
              • Grevilleas, Banksias and Allies - Subfamily: Grevilleoidea >
                • Banksias - Tribe: Banksieae
                • Firebushes and Allies - Tribe: Embothrieae
                • Macadamias and Allies - Tribe: Macadamieae
                • Honeysuckles and Allies - Tribe: Roupaleae
              • Primitive Proteas - Subfamily: Persoonioideae >
                • Geebungs and Allies - Tribe: Persoonieae
              • Singlet Proteas - Subfamily: Proteoideae >
                • Featherbushes and Conesticks - Tribe: Petrophileae
          • Buttercups, Poppies and Allies - Order: Ranunculales >
            • Buttercups - Family: Ranunculaceae
            • Moonseeds - Family: Menispermaceae
            • Poppies and Allies - Family: Papaveraceae
          • Roses, Figs, Elms and Allies - Order: Rosales >
            • Hemps and Allies - Family: Cannabaceae
            • Figs, Mulberries and Allies - Family: Moraceae
            • Buckthorns and Allies - Family: Rhamnaceae
            • Roses, Brambles and Allies - Family: Rosaceae
            • Elms and Allies - Family: Ulmaceae
            • Nettles and Allies - Family: Urticaceae
          • Sandalwoods, Mistletoes and Allies - Order: Santalales >
            • Showy Mistletoes - Family: Loranthaceae
            • Sandalwoods - Family: Santalaceae
          • Soapberries, Mahoganies and Allies - Order: Sapindales >
            • Cashews, Sumacs and Allies - Family: Anacardiaceae
            • Mahoganies and Allies - Family: Meliaceae
            • Rues, Citruses and Allies - Family: Rutaceae
            • Soapberries and Allies - Family: Sapindaceae
            • Quassias, Corkwoods and Allies - Family: Simaroubaceae
          • Saxifrages, Stonecrops and Allies - Order: Saxifragales >
            • Stonecrops and Allies - Family: Crassulaceae
          • Nightshades, Bindweeds and Allies - Order: Solanales >
            • Bindweeds, Ponysfoots and Allies - Family: Convolvulaceae
            • Nightshades, Tobacco and Allies - Family: Solanaceae
          • Grapes and Allies - Order: Vitales >
            • Grapes and Allies - Family: Vitaceae
    • Protozoans - Kingdom: Protozoa >
      • Slime Moulds - Phylum: Mycetozoa >
        • True Slime Moulds - Class: Myxomycetes
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